The final objective of this website’s is to provide a one-stop resource for concerned citizens who want to learn more about the shape of designer extinction.
Why? Because it’s time more concerned citizens made the move from half-hearted Environmentalism to full-on Gaianism. A Gaianist is one who not only discerns the shape of right relation with the living world, but attempts to embody that relation in one’s own day-to-day. Deep ecologists are Gaianists, so are members of the Extinction Rebellion. The CEOs of most large environmental organizations are not Gaianists – not if they believe going greem means swapping out from fuels for “clean energy” without at the same time clawing back on their own typically exorbitant lifestyles. That is Environmentalism, not Gaianism.
How? As with Charles Darwin and his barnacles, as with E.O. Wilson and his ants, as with David Suzuki and his fruit flies, so with Gaianism: one begins by loving and learning about some part of the living world, and then moves out from there. Deep-Snow Caribou shed illuminating light on the structure of our current social order. If only for this reason, they make a perfect entry point to apprenticeship. Hence this bibliography.
PAPERS & REPORTS PERTAINING TO WOODLAND CARIBOU, WITH SPECIAL REFERENCE TO DEEP-SNOW MOUNTAIN CARIBOU, WOLVES, OLDGROWTH INLAND RAINFORESTS & ARBOREAL HAIR LICHENS
NOTE: This webpage is under construction. More to come. Stay tuned.
CURRENT TO 31 JULY 2020
Adams, L.G. 2003. Marrow fat deposition and skeletal growth in caribou calves. Journal of Wildlife Management 67:20–24.
Adams, L.G. 2005. Effects of maternal characteristics and climatic variation on birth masses of Alaskan caribou. Journal of Mammalogy 86:506–513.
Aagnes, T.H., W. Sormo and S.D. Mathiesen. 1995. Ruminal microbial digestion in free-living, in captive lichen-fed, and in starved reindeer (Rangifer tranadus tarandus) in Winter. Applied and Environmental Microbiology Feb 1995: 583–591.
Andrews, W.J. 2017. Species at Risk Act, S.C. 2002, c. 29 Application to The Honourable Catherine McKenna, Minister of Environment and Climate Change and to Governor in Council for an Emergency Order under section 80 to Protect Matrix Range Critical Habitat of the Wells Gray-Thompson Local Population Unit of the Woodland Caribou, Southern Mountain Caribou population (Rangifer tarandus caribou) Against Timber Harvesting and Related Road-building on Provincial Crown Land. 68 pages + appendices. Available here. (Accessed August 2018.)
Anonymous. 2017. Wolf Management Plan for Caribou: Summary Year Three – 2016–17 FINAL – August 28 2017. Available here. (Accessed July 2018.)
Antifeau, T.D. 1987. The significance of snow and arboreal lichens in Wells Gray Provincial Park with special reference to their importance to mountain caribou (Rangifer tarandus caribou) in the North Thompson watershed of British Columbia. M.Sc. thesis, University of British Columbia. 142 pages.
Apps, C. and B. McLellan. 2006. Factors influencing the dispersion and fragmentation of endangered mountain caribou populations. Biological Conservation 130: 84–97. Available here.
Apps, C.D., B.N. McLellan, T.A. Kinley and J.P Flaa. 2001. Scale-dependent habitat selection by mountain caribou, Columbia Mountains, British Columbia. Journal of Wildlife Management 65: 65–77.
Apps, C.D., B.N. McLellan, T. Kinley, R. Serrouya, D. Seip and H. Wittmer. 2013. Spatial factors related to mortality and population decline of endangered mountain caribou. Journal of Wildlife Management 77: 1409–1419.
Armleder, H.M. and S.K. Stevenson. 1996. Using alternative silvicultural systems to integrate Mountain Caribou and timber management in British Columbia. Rangifer, Special Issue 9: 141–147.
Arsenault, A. and T. Goward. 2000. Ecological characteristics of inland rain forests. At risk: proceedings of a conference on the biology and management of species and habitats at risk, Kamloops, B.C., 15–19 February, 1999, Vol. 1 (ed. by L.M. Darling), pp. 437–439. BC Ministry of Environment, Lands and Parks, Victoria, BC and University College of the Cariboo, Kamloops, BC.
Auditor General of British Columbia. 2015. Managing the cumulative effects of natural resource development in British Columbia. Auditor General of British Columbia, Victoria. Available here. (Accessed October 2018.)
BC Ministry of Agriculture and Lands, Integrated Land Management Bureau. 2007. Mountain Caribou Recovery Implementation Plan, comprised of: News Release, Backgrounder, and Map. 16 October 2007. Available at xxx. (Accessed xxx)
BC Ministry of Environment. 2009. Mountain Caribou Recovery Implementation Plan: Update to the Mountain Caribou Progress Board, February 2009. Available here. (Accessed October 2018.)
Bergerud, A.T. 1974. Decline of caribou in North America following settlement. Journal of Wildlife Management 38: 757–770.
Bergerud, A.T. 1996. Evolving perspectives on caribou population dynamics, have we got it right yet? Rangifer Special Issue 9: 95–116.
Bergerud, A.T. 2000. Caribou. Chapter 11, pages 658–693 in Ecology and Management of Large Mammals in North America. S. Demarais and P.R. Krausmann (eds.). Prentice Hall, New Jersey.
Bergerud, A.T. and J.P. Elliot. 1986. Dynamics of caribou and wolves in northern British Columbia. Can. J. Zool. 64: 1515–1529.
Bird, C., R. Gill, R. Serrouya and H. van Oort. 2015–16. Lake Revelstoke wolf census. BC Ministry of Forests, Lands, and Natural Resource Operations, Nelson, BC. Available here. (Accessed August 2018).
Boan, J.J., J.R. Malcolm, M.D. Vanier, D.L. Euler and F.M. Moola. 2018. From climate to caribou: how manufacturing of uncertainty is affecting wildlife management. Wildlife Society Bulletin 42: 366–381.
Boutin, S. and E. Merrill. 2016. A review of population-based management of Southern Mountain caribou in BC. Report to Columbia Mountains Institute. March 2016. Available at xxx (Accessed October 2018)
Brook, R.K., M. Cattet, C.T. Darimont, P.C. Paquet and G. Proulx. 2015. Maintaining ethical standards during conservation crises. Canadian wildlife biology and management 4: 72–79.
Brockman, C.J., W.B. Collins, J.M. Welker, D.E. Spalinger and B.W. Dale. 2017. Determining kill rates of ungulate calves by brown bears using neck-mounted cameras. Wildlife Society Bulletin 41: 88–97.
Brook, R.K., M. Cattet, C.T. Darimont, P.C. Paquet and G. Proulx. 2015. Maintaining ethical standards during conservation crises. Canadian wildlife biology and management 4: 72–79.
Brown, G.S., L. Landriault, D.J.H. Sleep and F.F. Mallory. 2007. Comments arising from a paper by Wittmer et al.: hypothesis testing for top-down and bottom-up effects in woodland caribou populations dynamics. Oecologia 154: 485–492.
Brown, G. and F. Mallory. 2007. A review of ungulate nutrition and the role of top-down and bottom-up forces in woodland caribou population dynamics. National Council for Air and Stream Improvement. Technical Bulletin No. 934: 1–94.
Cameron, R.D., W.T. Smith, S.G. Fancy, K.L. Gerhart and R G. White. 1993. Calving success of female caribou in relation to body weight. Canadian Journal of Zoology 71: 480–486.
Campbell, J. and D.S. Coxson. 2003. Canopy microclimate and arboreal lichen loading in subalpine spruce-fir forest. Canadian Journal of Botany 79: 537–555.
Campbell, J., S.K. Stevenson and D.S. Coxson. 1999. Estimating epiphyte abundance in high elevation forests of Northern British Columbia. Selbyana 20: 261–267.
Ceballos, G., P.R. Ehrlich, A.B. Barnosky, A. Garcia, R.M. Pringle and T.M. Palmer. 2015. Accelerated modern human-induced species losses: entering the sixth mass extinction. Scientific Advances 1:e1400253.
Ceballos, G., P.R. Ehrlich and R. Dirzo. 2017. Biological annihilation via the ongoing sixth mass extinction signalled by vertebrate population losses and declines. Proceedings of the National Academy of Sciences USA. 114: E6089–E6096.
Cocklin, C., S. Parker and J. Hay. 1992. Notes on cumulative environmental change I: concepts and issues. Journal of Environmental Management 35: 31–49.
Collard, R.-C., J. Dempsey and M. Holmberg. 2019. Extirpation despite regulation? Environmental assessment and caribou. Conservation Science and Practice. 2020;2:e166. Available here. (Accessed July 2020.)
Cook, R.C., J.G. Cook, D.J. Vales, B.K. Johnson, S.M. McCorquodale, L.A. Shipley, R.A. Riggs, L.L. Irwin, S.L. Murphie, B.L. Murphie and K.A. Schoenecker. 2013. Regional and seasonal patterns of nutritional condition and reproduction in elk. Wildlife Monographs 184: 1–45.
Cook, J. and R. Cook. 2015. Nutritional condition of caribou in Northern British Columbia, 2012–2015 and Southern Northwest Territories. Annual Progress report, National Council for Air and Stream Improvement. 16 pages. (Not seen, cited in Boutin & Merrill 2016)
COSEWIC. 2002. COSEWIC assessment and update status report on the woodland caribou Rangifer tarandus caribou in Canada. Committee on the Status of Endangered Wildlife in Canada. Ottawa. xi + 98 pp. Available here. (Accessed October 2018.)
COSEWIC. 2011. Designatable Units for Caribou (Rangifer tarandus) in Canada. Committee on the Status of Endangered Wildlife in Canada. Ottawa. 88 pages. Available at xxx
COSEWIC. 2014. COSEWIC assessment and status report on the Caribou Rangifer tarandus, Northern Mountain population, Central Mountain population and Southern Mountain population in Canada. Committee on the Status of Endangered Wildlife in Canada. Ottawa. xxii + 113 pages. Available here. (Accessed October 2018.)
Cox, S. 2018a. All hype, no help: B.C. draws ire from scientists over caribou plan. The Narwhal 19 Oct 2018. Available here. (Accessed October 2018).
Cox, S. 2018b. BC approves 83 logging cutblocks in endangered caribou habitat in last six months. The Narwhal 23 Oct 2018. Available here. (Accessed October 2018.)
Cox, S. 2019a. “A sad day”: two more BC mountain caribou herds now locally extinct. The Narwhal 14 Jan 2019. Available here. (Accessed March 2019).
Cox, S. 2019b. BC approves 314 new cutblocks in endangered caribou habitat over last five months. The Narwhal 14 March 2019. Available here. (Accessed March 2019).
Cox, S. 2020a. ‘A dangerous road’: Coastal GasLink pays to kill wolves in endangered caribou habitat in B.C. interior. The Narwhal 13 April 2020. Available here.
Cox, S. 2020b. The complicated tale of why B.C. paid $2 million to shoot wolves in endangered caribou habitat this winter. The Narwhal 25 April 2020. Available here. (Accessed July 2020).
Coxson, D.S. and M. Coyle. 2003. Niche partitioning and photosynthetic response of lichens from subalpine spruce-fir forest in north-central British Columbia, Canada: the role of canopy microclimate. Lichenologist 35: 157–175.
Coxson, D.S. and M. Curteanu. 2002. Decomposition of hair lichens (Alectoria sarmentosa and Bryoria spp.) under snowpack in montane forest, Cariboo Mountains, British Columbia. Lichenologist 34: 395–402.
Coxson, D., T. Goward and J.R. Werner. 2020. The Inland Temperate Rainforest and Interior Wetbelt Biomes of Western North America. Pages 88–102 in M.I. Goldstein and D.A. DellaSala (Eds.), Encyclopedia of the World’s Biomes, vol. 3. Elsevier, pp. 88–102.
Ccc! Coxson, D.S. and Stevenson 2005 xxx
Della Sala, D., P. Alaback, L. Craighead, T. Goward, P. Paquet and T. Spribille. 2011. Chapter 3. Temperate and boreal rainforests of inland northwestern North America. Pp. 82–110 in D. Della Sala (ed.): Temperate and boreal rainforests of the world. Island Press, Washington.
DeMars, C. 2017. Recovery action plan for Southern Mountain caribou within the Kootenay Region. BC Ministry of Forests, Lands and Natural Resource Operations, Victoria. Not seen, cited in report available here. (Accessed October 2018)
Detrick, R.W.T. 1985. Effects of fire and logging on arboreal lichen availability to caribou. M.Sc. thesis, University of Idaho Graduate School. 49 pages.
Dickie, M., R. Serrouya, C. DeMars, J. Cranston and S. Boutin. 2017. Evaluating functional recovery of habitat for threatened woodland caribou. Ecosphere 8(9):e01936: 1–15.
Doak, D.F., J.A. Estes, B.S. Halpern, U. Jacob, D.R. Lindberg, J. Lovvorn, D.H. Monson, M.T. Tinker, T.M. Williams, J.T. Wootton, I. Carroll, M. Emmerson, F. Micheli and M. Novak. 2008. Understanding and Predicting Ecological Dynamics: Are Major Surprises Inevitable? Ecology 89: 952–961.
Dyer, S.J., J.P. O’Neill, S.M. Wasel and S. Boutin. 2001. Avoidance of industrial development by woodland caribou. The Journal of Wildlife Management 65: 531–542.
Edmonds, J. 1991. Status of woodland caribou in western North America. Rangifer, Special Issue No. 7: 91–107.
Edwards, R.Y. 1954. Fire and the decline of a mountain caribou herd. Journal of Wildlife Management 18: 521–526.
Edwards, R.Y. 1956. Snow depths and ungulate abundance in the mountains of western Canada. Journal of Wildlife Management 20: 159–168.
Edwards, R.Y. 1958. Land form and caribou distribution in British Columbia. Journal of Mammology 39: 408–412.
Edwards, R.Y. and R.W. Ritcey. 1959. Migrations of caribou in a mountainous area in Wells Gray Park, British Columbia. Canadian Field-Naturalist 73: 21–25.
Edwards, R.Y. and R.W. Ritcey. 1960. Foods of caribou in Wells Gray Park, British Columbia. Canadian Field-Naturalist 74: 3–7.
Edwards, R.Y., J. Soos and R.W. Ritcey. 1960. Quantitative observations of epidendric lichens used as food by caribou. Ecology 41: 425–431.
Elbroch, L.M., P.E. Lendrum, J. Newby, H. Quigley and D.J. Thompson. 2015. Recolonizing wolves influence the realized niche of resident cougars. Zoological Studies 54:41: 1–15.
Environment Canada. 2014. Recovery Strategy for the Woodland Caribou, Southern Mountain population (Rangifer tarandus caribou) in Canada. Species at Risk Act Recovery Strategy Series. Environment Canada, Ottawa. viii + 103 pages. Available here. (Accessed October 2018).
Environment and Climate Change Canada. 2018. Imminent Threat Assessment for Southern Mountain Caribou. 41 pages. Available here. (Accessed October 2018).
Esseen, P.-A., T. Olsson., D. Coxson and Y. Gauslaa. 2015. Morphology influences water storage in hair lichens from boreal forest canopies. Fungal Ecology 18: 26–35.
Festa-Bianchet, M., J.C. Ray, S. Boutin, S.D. Côté and A. Gunn. 2011. Conservation of caribou (Rangifer tarandus) in Canada: an uncertain future. Can. J. Zool. 89: 419–434.
Fisher, J.T. and L. Wilkinson. 2005. The response of mammals to forest fire and timber harvest in the North American boreal forest. Mammal Review 25: 51–18.
Flaa, J.P. and B.N. McLellan. 2000. Population characteristics of the Lake Revelstoke caribou. Darling (ed.). Proceedings of a conference on the biology and management of species and habitats at risk, Kamloops, B.C., 15–19 Feb., 1999. B.C. Ministry of Environment, Lands and Parks, Victoria, B.C. and University College of the Cariboo, Kamloops, B.C.
Forest Practices Board. 2011. Cumulative effects: from assessment towards management, Special Report, FPB/SR/39, March 2011. Available at xxx (Accessed October 2018)
Forest Practices Board. 2015a. Timber Removal in Mountain Caribou Habitat, Special Investigation, FPB/SIR/45, December 2015. Available here. (Accessed October 2018).
Forest Practices Board. 2015b. Access Management and Resource Roads: 2015 update. Special Report, FPB/SR/49, April 2015. Available here. (Accessed October 2018).
Freddy, D.J. 1974. Status and management of the Selkirk caribou herd, 1973. M.Sc. Thesis. University of Idaho. 132 pages.
Freeman, N. 2008. Motorized backcountry recreation and stress response in Mountain Caribou (Rangifer tarandus caribou). M.Sc. Thesis, University of British Columbia, Vancouver. 75 pages.
Furk, K., M. McLellan and R. Serrouya. 2008. Condition indices of caribou (Rangifer tarandus caribou) found at mortality sites in the North Columbia Mountains of British Columbia. Unpublished report to BC Ministry of Forests and Range, March 2008. 18 pages.
Furk, K. and R. Serrouya. 2016. Revelstoke Caribou Maternity Pen Project Annual Report, Year 2, April 1st, 2015-March 31st, 2016. Unpublished report, Revelstoke Caribou Rearing in the Wild, Revelstoke, BC. Available here. (Accessed August 2018).
Furk, K. and R. Serrouya. 2017. Revelstoke Caribou Maternity Pen Project Annual Report, Year 2, April 1st, 2017-March 31st, 2017. Unpublished report, Revelstoke Caribou Rearing in the Wild, Revelstoke, BC. 21 pages. Available here. (Accessed August 2018).
Furk, K., R. Serrouya and Cory Legebokow. 2011. Population censuses of mountain caribou in the North Columbia Mountains: March 2011 – Columbia North, Columbia South, Frisby-Queest and Monashee South subpopulations. Unpublished report to BC Ministry of Forests and Range, March 2011. 20 pages.
Garrott, R.A., J.E. Bruggeman, M.S. Becker, S.T. Kalinowski and P.J. White. 2007. Evaluating prey switching in wolf-ungulate systems. Ecological Applications 17: 1588–1597.
Gerhart, K.L., D.E. Russell, D. Van de Wetering, R.G. White and R.D. Cameron. 1997. Pregnancy of adult caribou (Rangifer tarandus): evidence for lactational infertility. Journal of Zoology 242: 17–30.
Gillis, D. 2017a. BC Liberals grant major political donor permission to log endangered caribou habitat. The Narwhal 3 April 2017. Available here. (Accessed October 2018).
Gillis, D. 2017b. Endangered Caribou habitat clearcut during B.C. election uncertainty. The Narwhal 26 July 2017. Available here. (Accessed October 2018).
Government of Canada. 2017. Canada-British Columbia Southern Mountain Caribou (Central Group) Protection Study. February 2017. 113 pages. Available here. (Accessed October 2018).
Goward, T. 1994. Notes on oldgrowth-dependent epiphytic macrolichens in the humid oldgrowth forests in inland British Columbia, Canada. Acta Botanica Fennica 150: 31–38.
Goward, T. 1998. Observations on the ecology of the lichen genus Bryoria in high elevation conifer forests. Canadian Field-Naturalist 112: 496–501.
Goward, T. 2002. Hair lichens, snowpack variation and the fate of the mountain caribou: the LSC hypothesis. Abstract. Pages 41–42 in Mountain Caribou in 21st century ecosystems. Revelstoke. Columbia Mountains Institute of Applied Ecology. Available here. (Accessed July 2018).
Goward, T. 2003a. On the vertical zonation of hair Lichens (Bryoria) in the canopies of high-elevation oldgrowth conifer forests. The Canadian Field-Naturalist 114: 39–43.
Goward, T. 2003b. On the dispersal of hair lichens (Bryoria) in high-elevation oldgrowth conifer forests. The Canadian Field-Naturalist 117: 44–48.
Goward, T. and J. Campbell. 2005. Arboreal hair lichens in a young, unmanaged, mid-elevation conifer stand, with implications for Mountain Caribou. The Bryologist 108: 427–434.
Goward, T. and J. Pojar. 1998. Antique forests and epiphytic macrolichens in the Kispiox Valley. B.C. Forest Service Extension Note 33: 1–1l.
Goward, T. and A. Arsenault. 2000. Inland oldgrowth rainforests: safe havens for rare lichens? Pages 759–766 in L. Darling (ed.). Proceedings of a conference on the biology and management of species and habitats at risk, Kamloops, B.C., 15–19 Feb., 1999. B.C. Ministry of Environment, Lands and Parks, Victoria, B.C. and University College of the Cariboo, Kamloops, B.C.*
Hansen, M.J., Franklin, S.E., Woudsma, C.G. and M. Peterson. 2001. Caribou habitat mapping and fragmentation analysis using Landsat MSS, TM, and GIS data in the North Columbia Mountains, British Columbia, Canada. Remote sensing of Environment 77: 50–65.
Gustine, D.D., P.S. Barboza, L.G. Adams and N.B. Wolf. 2014. Environmental and physiological influences to isotopic ratios of N and protein status in a montane ungulate in winter. PloS One 9:e103471.
Harding, L. 1975. Our Mountain Caribou… endangered species? BC Outdoors 31: 24–31.
Harding, L.E. 2017. Request for an Emergency Order under the Species at Risk Act, S.C. 2002, c. 29, section 80, to Protect Matrix Range Critical Habitat of the Central Selkirk Local Population Unit of the Woodland Caribou, Southern Mountain Caribou population (Rangifer tarandus caribou) Against Timber Harvesting and Related Road-building on Provincial Crown Land. Coquitlam, B.C., Letter sent to The Hon. Katherine McKenna, Minister of Environment and Climate Change, Government of Canada, 14 April 2017. 14 pages.
Harding, L., M. Bourbonnais, A.T. Cook, T. Spribille, V. Wagner and C. Darimont. 2020. No statistical support for wolf control and maternal penning as conservation measures for endangered mountain caribou. Biodiversity and Conservation. Available here.
Hatter, I.W. 2000. Components of a Mountain Caribou recovery strategy for British Columbia. Pp. 631–635 in L.M. Darling, ed. Proceedings of a Conference on the Biology and Management of Species and Habitats at Risk, Kamloops, BC, 15–19 Feb. 1999. Vol. 2. BC Ministry of Environment, Lands and Parks, Victoria, BC and University College of the Cariboo, Kamloops, BC.
Hatter, I. 2006. Mountain caribou 2006 survey results, subpopulation trends and extinction risk. Draft for Technical Review. BC Ministry of Environment. Victoria, B.C. 19 pages.
HCMRIG (Hart and Cariboo Mountains Recovery Implementation Group) 2005. Recovery implementation plan for threatened woodland caribou (Rangifer tarandus caribou) in the Hart and Cariboo Mountains recovery area, British Colum¬bia, Canada. Unpublished report. Available here. (Accessed October 2018).
Heard, D.C. and K.L. Vagt. 1998. Caribou in British Columbia: a 1996 status report. Rangifer Spec. Issue, 10: 117–123.
Hebblewhite, M., C. White and M. Musiani. 2010. Revisiting extinction in National Parks: mountain caribou in Banff. Conservation Biology 24: 341–344.
Hebblewhite, M. 2017. Billion-dollar boreal woodland caribou and the biodiversity impacts of the global oil and gas industry. Biological Conservation 206: 102–111.
Hegel, T.M., A. Mysterud, F. Huettmann and N.C. Stenseth. 2010a. Interacting effects of wolves and climate on recruitment in a northern mountain caribou population. Oikos 119: 1453–1461.
Hegel, T.M., A. Mysterud, T. Ergon, L.E. Loe, F. Huettmann and N.C. Stenseth. 2010b. Seasonal effects of Pacific-based climate on recruitment in a predator-limited large herbivore. Journal of Animal Ecology 79: 471–482.
Hervieux, D., M. Hebblewhite, D. Stepnisky, M. Bacon and S. Boutin. 2014. Managing wolves (Canis lupus) to recover threatened woodland caribou (Rangifer tarandus caribou) in Alberta. Canadian Journal of Zoology 92: 1029–1037.
Holt, R.D. 1977. Predation, apparent competition, and the structure of prey communities. Theoretical Population Biology 11: 197–229.
Holt, R. 1984. Spatial Heterogeneity, Indirect Interactions, and the Coexistence of Prey Species. The American Naturalist 124: 377–406.
Holt, R.D. 2008. Theoretical perspectives on resource pulses. Ecology 89: 671–681.
Hooge, J., B. McLellan and J. Flaa. 2004. 2004 population census of Mountain Caribou in the Columbia Forest District. Unpublished report to B.C. Ministry of Forests, Columbia Forest District, Revelstoke. 11 pages. Available at xxx (Accessed July 2018 xxx)
Hornseth, M.L. and R.S. Rempel. 2015. Seasonal resource selection of woodland caribou (Rangifer tarandus caribou) across a gradient of anthropogenic disturbance. Canadian Journal of Zoology 94: 79–93.
Hsieh, W.W. and B. Tang. 2001. Interannual variability of accumulated snow in the Columbia basin, British Columbia. Water Resources Research 37: 1753–1759.
Hunter, M.D. and P.W. Price. 1992. Playing chutes and ladders: heterogeneity and the relative roles of bottom-up and top-down forces in natural communities. Ecology 73: 724–732.
Janzen, D.H. 1983. No park is an island: increase in interference from outside as park size decreases. Oikos 41: 402–410.
Johnson, C.J., L.P.W. Ehlers and D.R. Seip. 2015. Witnessing extinction – Cumulative impacts across landscapes and the future loss of an evolutionarily significant unit of woodland caribou in Canada. Biological Conservation 186: 176–186.
Johnson, C.J., M.A. Mumma and M-H. St-Laurent. 2019. Modeling multispecies predator–prey dynamics: predicting the outcomes of conservation actions for woodland caribou. Ecosphere 10(3):e02622. 10.1002/ecs2.2622
Johnson, C., K. Parker, D. Heard and M. Gillingham. 2002. A multiscale behavioural approach to understanding the movements of woodland caribou. Ecological Applications 12: 1840–1860.
Johnson, D.R. 1976. Mountain Caribou: threats to survival in the Kootenay Pass region, British Columbia. Northwest Science 50: 97–101.
Johnson, D.R. and D.R. Miller. 1979. Observations on reproduction of Mountain Caribou. Northwest Science 53: 114–117.
Kerr, J.T. and I. Deguise. 2004. Habitat loss and the limits to endangered species recovery. Ecology Letters 7: 1163–1169.
Kinley, T.A. 1999. Wildlife in British Columbia at risk: Mountain Caribou. BC Ministry of Environment, Lands and Parks, Wildlife Branch, Victoria, BC. 6 pages.
Kinley, T.A. and C.D. Apps. 2001. Mortality patterns in a herd of endangered mountain caribou. Wildlife Society Bulletin 29: 158–164.
Kinley, T.A., T. Goward, B.N. McLellan and R. Serrouya. 2007. The influence of variable snowpacks on habitat use by Mountain Caribou. Rangifer, Special Issue 17: 93–102.
Klein, D.R. 1991. Limiting factors in caribou population ecology. Rangifer Special Issue 7: 30–35.
Kolbert, E. 2014. The sixth extinction: an unnatural history. Henry Holt and Company, New York. 319 pages.
Krausman, P.R. and L.K. Harris. 2011. Cumulative effects in wildlife management: impact mitigation. CRC Press, Taylor and Francis Group, Boca Raton, Florida.
Krebs, C.J. 2002. Beyond population regulation and limitation. Wildlife Research 29: 1–10.
Kuussaari, M., Bommarco, R., Heikkinen, R.K., Helm, A., Krauss, J., Lindborg, R., Öckinger, E., Pärtel, Pino, M.J., Rodà, F., Stefanescu, C., Teder, T., Zobel, M. and Steffan-Dewenter, I. 2009. Extinction debt: a challenge for biodiversity conservation. Trends in Ecology & Evolution 24: 564–571.
Lande, R. 1993. Risks of population extinction from demographic and environmental stochasticity and random catastrophes. American Naturalist 142: 911–927.
Lavoie, J. 2018a. Seeking the science behind B.C.’s wolf cull. The Narwhal 5 April 2018. Available here. (Accessed October 2018).
Lavoie, J. 2018b. Caribou on the brink: B.C. herd reduced to three females points to failure to protect endangered species. The Narwhal 18 April 2018. Available here. (Accessed October 2018).
Lavoie, J. 2018c. Minister’s inaction on B.C.’s endangered caribou ‘egregious’” federal court judge. The Narwhal 17 May 2018. Available here. (Accessed October 2018).
Layser, E.F. 1974. A review of the mountain caribou of northeastern Washington and adjacent northern Idaho. Journal of the Idaho Academy of Science, Special Research Issue 3: 1–63.
Leech, H., D.E. Jelinski, L. DeGroot and G.W. Kuzyk. 2017. The temporal niche and seasonal differences in predation risk to translocated and resident caribou (Rangifer tarandus caribou). Canadian Journal of Zoology 25: 809–820.
Leo, J.Y.H. 1994. Effects of growing elevation and host tree species on the crude protein and gross energy of arboreal lichens (Alectoria and Bryoria spp.) in the winter diet of mountain caribou (Rangifer tarandus caribou). Bachelor of Science thesis, Department of Animal Science, UBC, May 1994 38 pages. [paper copy only]
Lindenmayer, D.B., M.L. Hunter, P.J. Burton and P. Gibbons. 2009. Effects of logging on fire regimes in moist forests. Conservation Letters 2: 271–277.
Lindenmayer, D.B. and J. Fischer. 2006. Habitat fragmentation and landscape change: an ecological and conservation synthesis. Island Press, Washington, DC.
McLellan, B.N. 2010. An historic perspective of mountain caribou distribution and abundance. BC Ministry of Forests, Research Branch, Victoria, British Columbia. Unpublished manuscript. Available here. (Accessed October 2018).
McLellan, B.N., J. Flaa and J. Woods. 1994. Mountain caribou censuses in the North Columbia Mountains. Unpublished report to B.C. Ministry of Forests and Canadian Parks Service, Revelstoke. 15 pages. Available here. (Accessed October 2018).
McLellan, B., R. Serrouya and J. Flaa. 2006. Population censuses of caribou the North Columbia Mountains. Unpublished report to B.C. Ministry of Forests, Revelstoke. 13 pages. Available here. (Accessed October 2018).
McLellan, M., R. Serrouya, B.N. McLellan, K. Furk, D. Heard and H. Wittmer. 2012. Implications of body condition on the unsustainable predation rates of endangered mountain caribou. Oecologia 169: 853–860.
McLellan, B.N., R. Serrouya, H.U. Wittmer and S. Boutin. 2010. Predator-mediated Allee effects in multi-prey systems. Ecology 91: 286–292. Available here.
MCRIP Progress Board. 2012. Mountain Caribou Recovery Implementation Plan Progress Board: Annual Report on Activities and Accomplishments of the Mountain Caribou Recovery 2011–12, Unpublished report to BC Ministry of Environment. 27 pages. Available here. (Accessed July 2018).
MCST [Mountain Caribou Science Team]. 2005. Mountain Caribou in British Columbia: a situation analysis. Available at xxx (Accessed July 2018 xxx)
MCTAC [Mountain Caribou Technical Advisory Committee]. 2002. A strategy for the recovery of Mountain Caribou in British Columbia. Mountain Caribou Technical Advisory Committee. B.C. Ministry of Water, Land and Air Protection. Victoria. 73 pages. Available at xxx (Accessed July 2018 xxx)
MFLNRO LNG. 2015. LNG Caribou Program: A plan to support the conservation and management of Northern and Mountain Caribou populations and habitat potentially affected by LNG pipeline projects, North Area. BC Ministry of Forests, Lands and Natural Resource Operations, Victoria. Not seen, cited in report available here. (Accessed October 2018).
Meserve, P.L., D.A. Kelt, W.B. Milstead and J.R. Gutiérrez. 2003. Thirteen years of shifting top-down and bottom-up control. BioScience 53: 633–646.
Messier, C., K. Puettmann, R. Chazdon, K.P. Anderson, V.A. Angers, L. Brotons, E. Filotas, R. Tittler, L. Parrott and S.A. Levin. 2015. From management to stewardship: viewing forests as complex adaptive systems in an uncertain world. Conservation Letters 8: 368–77.
Messier, C., K. Puettmann, E. Filotas and D. Coates. 2016. Dealing with non-linearity and uncertainty in forest management. Current Forestry Reports 2: 150–161.
Miller, F.L. and A. Gunn. 2003. Catastrophic Die-Off of Peary Caribou on the Western Queen Elizabeth Islands, Canadian High Arctic. Arctic 56: 381–390.
Mosnier, A., D. Boisjoly, R. Courtois and J.-P. Ouellet. 2008. Extensive predator space use can limit the efficacy of a control program. Journal of Wildlife Management 72: 483–491. doi:10.2193/2006 xxx
MCST [Mountain Caribou Science Team]. 2005. Mountain Caribou in British Columbia: a situation analysis. Unpublished report to BC Ministry of Environment, Victoria. Available here. (Accessed October 2018).
Newmark, W.D. 2002. Extinction of mammal populations in western North America. Conservation Biology 9: 512–526.
Nikiforuk, A. 2017. Another Wild West Show? BC’s regulatory experiment with professional reliance. The Tyee 21 February 2017. Available here. (Accessed July 2018).
van_Oort, H., B.N. McLellan and R. Serrouya. 2011. Fragmentation, dispersal and metapopulation function in remnant populations of endangered mountain caribou. Animal Conservation 14: 215–224.
Palm, E.C., S. Fluker, H.K. Nesbitt, A.L. Jacob and M. Hebblewhite. 2020. The long road to protecting critical habitat for species at risk: the case of southern mountain woodland caribou. Conservation Science and Practice. 2020;e219. Available here. (Accessed July 2020).
Parker, K.L., P.S. Barboza and M.P. Gillingham. 2009. Nutrition integrates environmental responses of ungulates. Functional Ecology 23: 57–69.
Parker, K.L., P.S. Barboza and T.R. Stephenson. 2005. Protein conservation in female caribou (Rangifer tarandus): effects of decreasing diet quality during winter. Journal of Mammalogy 86: 610–622.
Peters, W. 2010. Resource selection and abundance estimation of moose: implications for caribou recovery in a human altered landscape. M.Sc. Thesis. University of Montana, Missoula, Montana, USA. 104p.
Pinney, N.H., K.A. Solhaug and Y. Gauslaa. 2018. Rapid resurrection of chlorolichens in humid air: specific thallus mass drives rehydration and reactivation kinetics. Environmental and Experimental Botany 148: 184–191.
Polfus, J.L., M. Hebblewhite and K. Heinemeyer. 2011. Identifying indirect habitat loss and avoidance of human infrastructure by northern mountain woodland caribou. Biological Conservation 144: 2637–2646.
Xxx Potapov, P., M.C. Hansen, L. Laestadius, S. Turubanova, A. Yaroshenko, C. Thies, W. Smith, I. Zhuravleva, A. Komarova, S. Minnemeyer and E. Esipova. 2017. The last frontiers of wilderness: tracking loss of intact forest landscapes from 2000 to 2013. Science Advances 2017: 3:e1600821. Available here.
Potapov, P., M.C. Hansen, L. Laestadius, S. Turubanova, A. Yaroshenko, C. Thies, W. Smith, I. Zhuravleva, A. Komarova, S. Minnemeyer and E. Esipova. 2017. The last frontiers of wilderness: tracking loss of intact forest landscapes from 2000 to 2013. Science Advances 2017: 3:e1600821. Available here.
Power, M.E. 1992. Top-down and bottom-up forces in food webs: Do plants have primacy? Ecology 73: 733–746.
Ray xxx and xxx Johnson. 2018. Response to BC Caribou Recovery Program Discussion Paper, 14 June 2018. Available here. (Accessed July 2018).
Revelstoke Caribou Rearing in the Wild Society. 2018. Revelstoke Maternal Penning Project Summer Update 2018. RCRW Factsheet 7, July 2018. Available here. (Accessed October 2018).
Rhoades, F.M. 1995. Nonvascular epiphytes in forest canopies: worldwide distribution, abundance, and ecological roles. Pages 353–408 in Lowman, M.D. and N. Nadkarni (eds.) Forest Canopies. Academic Press. New York.
Reimers, E., S. Eftestøl and J.E. Colman. 2003. Behavior responses of wild reindeer to direct provocation by a snowmobile or skier. The Journal of Wildlife Management 67: 747–754.
Ritcey, R.W. 1970. Caribou management and inventory in Management Area 12. Unpublished report, BC Ministry of Environment, Kamloops, BC. 6 pages.
Ritcey, R.W. 1974. Caribou and forest management in British Columbia. Unpublished report, B.C. Fish and Wildlife Branch, Kamloops. 10 pages.
Ritcey, R.W. 1976. Caribou management plan, Region III. B.C. Fish and Wildlife Branch, Kamloops. 15 pages.
Ritcey, R.W. 1981. Woodland caribou in the Thompson Nicola resource region. Unpublished report, B.C. Ministry of Environment. Kamloops. 18 pages.
Robbins, C.T. 1993. Wildlife feeding and nutrition. Second Edition. Academic Press, New York, New York. 352 pages.
Robbins, J. 2018. Gray Ghosts, the last caribou in the lower 48 states, are ‘functionally extinct.’ New York Times 14 April 2018. Available here. (Accessed July 2018).
Robinson, H., M. Hebblewhite, N. DeCesare, J. Whittington, L. Neufeld, M. Bradley and M. Musiani. 2012. The effect of fire on spatial separation between wolves and caribou. Rangifer Special Issue No. 20: 277–294.
Rominger, E.M. and J.L. Oldemeyer. 1989. Early-winter habitat of woodland caribou, Selkirk Mountains, British Columbia. Journal of Wildlife Management 53: 238–243.
Rominger, E.M. and J.L. Oldemeyer. 1990. Early-winter diet of woodland caribou in relation to snow accumulation. Canadian Journal of Zoology 68: 2691–2694.
Rominger, E.M. and C.T. Robbins. 1996. Generic preference and in-vivo digestibility of alectorioid arboreal lichens by woodland caribou. Rangifer Special Issue 9: 379–380.
Rominger, E.M., C.T. Robbins and M.A. Evans. 1996. Late-winter foraging dynamics of woodland caribou. Journal of Wildlife Management 60: 719–728.
Rominger, E.M., M.A. Evans, C.T. Robbins and D.J. Pierce. 2000. Autumn foraging dynamics of woodland caribou in experimentally manipulated habitats, northeastern Washington, USA. Journal of Wildlife Management 64: 160–167.
Schroeder, G.J. 1972. Quantification of the epiphytic lichen forage in the Selkirk caribou range: a progress report. Report presented to the Caribou Steering Committee, Dec. 1972, Spokane, Washington. 29 pages (not seen, cited in Stevenson et al. 1979).
Seip, D.R. 1989. Caribou-moose-wolf interactions in central British Columbia. Pages 57–69 in D. Seip, S. Pettigrew and R. Archibald (eds.). Wolf-prey dynamics and management. B.C. Ministry of Environment, Wildlife Working Report No. WR-40, Victoria.
Seip, D.R. 1990. Ecology of Woodland Caribou in Wells Gray Provincial Park. B.C. Ministry of Environment. Wildlife Bulletin No. B-68. 43 pp.
Seip, D.R. 1991. Predation and caribou populations. Rangifer Special Issue 7: 46–52.
Seip, D.R. 1992. Factors limiting woodland caribou populations and their interrelationships with wolves and moose in southeastern British Columbia. Canadian Journal of Zoology 71: 1494–1503.
Seip, D.R. 1998. Ecosystem management and the conservation of caribou habitat in British Columbia: what does it mean to put caribou knowledge into an ecosystem context? Rangifer Special Issue 0: 203–211.
Seip, D.R. 2008. Mountain caribou interactions with wolves and moose in central British Columbia. Alces 44: 1–5.
Seip, D.R. and D.B. Cichowski. 1996. Population ecology of caribou in British Columbia. Rangifer Special Issue 9: 73–80.
Seip, D., C. Johnson and G. Watts. 2007. Displacement of mountain caribou from winter habitat by snowmobiles. Journal of Wildlife Management 71: 1539–1544.
Seip, D.R. and B. McLellan. 2008. Mountain Caribou. Pages 240–255 in M. Hummel and J.C. Ray. Caribou and the north: a shared future. Dundurn Press, Toronto. 288 pages.
Serrouya, R.D. 2013. An adaptive approach to endangered species recovery based on a management experiment: reducing moose to reduce apparent competition with woodland caribou. Ph.D. thesis, University of Alberta. 220 pages.
Serrouya, R., K. Furk and C. Legebokow. 2016. Population surveys of Columbia North and Columbia South mountain caribou, March 2016. 15 pages. Unpublished report. Available here. (Accessed October 2018).
Serrouya, R., S. Gilbert, R.S. McNay, B.N. McLellan, D.C. Heard, D.R. Seip and S. Boutin. 2017a. Comparing population growth rates between census and recruitment-mortality models. Journal of Wildlife Management 81: 297–305.
Serrouya, R., B.N. McLellan and J.P. Flaa. 2007. Scale-dependent microhabitat selection by threatened mountain caribou (Rangifer tarandus caribou) in cedar-hemlock forests during winter. Canadian Journal of Forestry Research 37: 1082–1092.
Serrouya, R., B.N. McLellan, H. van Oort, G. Mowat and S. Boutin. 2017b. Experimental moose reduction lowers wolf density and stops decline of endangered caribou. Peer Journal August, 29, 2017.
Serrouya, R., D. Paetkau, B.N. McLellan, S. Boutin, M. Campbell and D.A. Jenkins. 2012. Population size and major valleys explain microsatellite variation better than taxonomic units for caribou in western Canada. Molecular Ecology 21: 2588–2601.
Serrouya, R., D.R. Seip, D. Hervieux, B.N. McLellan, RS. McNay, R. Steenweg, D.C. Heard, M. Hebblewhite, M. Gillingham and S. Boutin. 2019. Saving endangered species using adaptive management. Proceedings of the National Academy of Sciences USA.1816923116. Available here.
Serrouya, R., Wittmann, M.J., B.N. McLellan, H.U. Wittmer and S. Boutin. 2015. Using predator‐prey theory to predict outcomes of broad scale experiments to reduce apparent competition. The American Naturalist 185, 665–679.
Serrouya, R. and B.N. McLellan. 2016. Next steps for Southern Mountain Caribou recovery in planning Unit 3A, the Revelstoke Shuswap Region. Unpublished report. Revelstoke. 11 pages. Available here. (Accessed July 2018).
Serrouya, R., B.N. McLellan, H. van Oort, G. Mowat and S. Boutin. 2017. Experimental moose reduction lowers wolf density and stops decline of endangered caribou. Peer J, August, 29, 2017.
Simpson, K., J.P. Kelsall and M. Leung. 1996. Integrated management of mountain caribou and forestry in southern British Columbia. Rangifer, Special Issue 9: 153–158.
Sinclair, A.R.E. and C.J. Krebs. 2002. Complex numerical responses to top-down and bottom-up processes in vertebrate populations. Philosophical Transactions of the Royal Society 357: 1221–1231.
Sinclair, A.R.E., R.P. Pech, C.R. Dickman, D. Hik, P. Mahon and A.E. Newsome. 1998. Predicting effects of predation on conservation of endangered prey. Conservation Biology 12: 567–575.
Smith, K.G., E.J. Ficht, D. Hobson, T. Sorenson and D. Hervieux. 2000. Winter distribution of woodland caribou in relation to clear-cut logging in west-central Alberta. Canadian Journal of Zoology 78: 1433–1440.
Sorensen, T., P.S. McLoughlin, D. Hervieux, E. Dzus, J. Nolan, B. Wynes and S. Boutin. 2008. Determining sustainable levels of cumulative effects for boreal caribou. Journal of Wildlife Management 72: 900–905.
Soulé, M.E. and J. Terborgh. 1999. Conserving nature at regional and continental scales—a scientific program for North America. Bioscience 49: 809–817.
Spalding, D.J. 1990. The early history of moose (Alces alces): distribution and relative abundance in British Columbia. Contr. to Nat. Sci. #11. Royal B.C. Mus. Victoria, BC. 12pp.
Spalding, D.J. 2000. The early history of Woodland Caribou (Rangifer tarandus caribou) in British Columbia. British Columbia Ministry of Environment, Lands and Parks, Wildlife Branch, Victoria, British Columbia. Wildlife Bulletin Number B-100. 61 pages.
Steenweg, R.W. 2011. Interactions of wolves, mountain caribou and an increased moose-hunting quota – primary prey management as an approach to caribou recovery. MSc thesis, University of Northern British Columbia, Prince George. 144 pages.
Stevenson, S.K. 1979. Effects of selective logging on arboreal lichens used by Selkirk caribou. BC Fish and Wildlife Report No. R-2. 77 pages. Available here. (Accessed October 2018).
Stevenson, S.K. 1991. Forestry and caribou in British Columbia. Rangifer, Special Issue 7: 124–129.
Stevenson, S. 2001. Arboreal lichens. Pages 60–65 in M.J. Jull and S.K. Stevenson. The Lucille Mountain Study. 8-year results of a silvicultural systems trial in the Engelmann Spruce-Subalpine Fir zone. BC Ministry of Forests Research Branch Working Paper 59. Victoria, BC. Available here. (Accessed February 2019).
Stevenson, S.K., H.M. Armleder, M.J. Jull, D.G. King, B.N. McLellan and D.S. Coxson. 2001. Mountain caribou in managed forests: recommendations for managers. Second edition. Wildlife Report No. R-26, Ministry of Environment, Lands and Parks, Victoria, British Columbia. 58 pages. Available here. (Accessed October 2018).
Stevenson, S.K., H. Armleder, A. Arsenault, D. Coxson, C. DeLong and M. Jull. 2011. Ecology, Conservation, and Management of British Columbia’s Inland Rainforest. UBC Press. Vancouver, B.C.
Stevenson, S.K., K.N. Child, G.S. Watts and E.L. Terry. 1991. The Mountain Caribou in Managed Forests Program: Integrating forestry and habitat management in British Columbia. Rangifer, Special Issue 7: 130–136.
Stevenson, S.K. and D.S. Coxson. 2003. Litterfall, growth, and turnover of arboreal lichens after partial cutting in an Engelmann spruce-subalpine fir forest in northcentral British Columbia. Canadian Journal of Forest Research 33: 2306–2320.
Stevenson, S. and D. Coxson. 2007. Arboreal forage lichens in partial cuts – a synthesis of research results from British Columbia, Canada. Rangifer, Special Issue 17: 155–165.
Stevenson, S. and D. Hatler. 1985. Woodland caribou and their habitat in southern and central British Columbia. B.C. Ministry of Forests, Victoria, BC. Land Management Report 23: 1–354. (Volume 1).
Stevenson, S. and D. Hatler. 1985. Woodland caribou and their habitat in southern and central British Columbia. B.C. Ministry of Forests, Victoria, B.C. Land Management Report No. 23 (Volume 1). 354 pages.
Stotyn, S.A. 2008. Ecological Interactions of Mountain Caribou, wolves and moose in the North Columbia Mountains, British Columbia. M. Sc. Thesis, University of Alberta, Edmonton. 126 pages.
Suzuki, N. and K.L. Parker. 2019. Proactive conservation of high-value habitat for woodland caribou and grizzly bears in the boreal zone of British Columbia, Canada. Biological Conservation 230: 91–103.
Telfer, E.S. and J.P. Kelsall. 1984. Adaptations of some large North American mammals for survival in snow. Ecology 65: 1828–1834.
Terry, E.L., B.N. McLellan and G.S. Watts. 2000. Winter habitat ecology of mountain caribou in relation to forest management. Journal of Applied Ecology 37: 589–602.
The Hon. Pat Bell. 2008. Fourth Session, 38th Parliament official report of debates of the BC legislative assembly, Thursday10 April 2008, Morning Sitting Volume 30, Number 4. Hansard, Victoria, BC. 33 pages.
Thompson, V. and L. Harding. 2018. Application to The Honourable Catherine McKenna, Minister of Environment and Climate Change and to Governor in Council for an Emergency Order under section 80 to Protect Critical Habitat of the Revelstoke Shuswap Local Planning Unit of the Woodland Caribou, Southern Mountain Caribou population (Rangifer tarandus caribou) Against Timber Harvesting and Related Road Building on Provincial Crown Land. 73 pages + appendices.
Valhalla Wilderness Watch. 2007. Mountain Caribou greenwash: a critique of the BC government’s plan to protect the endangered Mountain Caribou. New Denver, BC. 23 pages. Available here. (Accessed July 2018).
Valhalla Wilderness Society. 2017. Petition for an emergency order under section 80(2) of the Species at Risk Act to immediately stop logging and motorized winter recreation in critical winter habitats of the Southern Mountain Caribou subpopulation of woodland caribou (Rangifer tarandus caribou). 20 November 2017/ 14 December 2017. Available here. (Accessed October 2018).
Valhalla Wilderness Watch. 2015. Press release: BC’s Mountain Caribou Recovery Plan is failing disastrously. New Denver, BC. 3 pages. Avaliable here. (Accessed July 2018).
Vors, L.S. and M.S. Boyce. 2009. Global declines of caribou and reindeer. Global Change Biology 15: 2626–2633.
Watson, J.E., T. Evans, O. Venter, B. Williams et al. 2018. The exceptional value of intact forest ecosystems. Nature Ecology and Evolution. Available here. (Accessed July 2018).
Whittington, J., M. Hebblewhite, N.J. DeCesare, L. Neufeld, M. Bradley, J. Wilmshurst and M. Musiani. 2011. Caribou encounters with wolves increase near roads and trails: a time-to-event approach. Journal of Applied Ecology 48: 1535–1542.
Wiles, G.J. 2017. Periodic status review for the woodland caribou in Washington. Washington Department of Fish and Wildlife, Olympia, Washington. 24 pages.
Wilson, S.F. and J.B. Nyberg. 2009. A Proposed Monitoring and Adaptive Management Strategy for Mountain Caribou Recovery Implementation. Unpublished report to BC Ministry of Environment, Victoria, B.C. 28 pages. Available here. (Accessed October 2018).
Wittmer, H. 2004. Mechanisms underlying the decline of mountain caribou (Rangifer tarandus caribou) in British Columbia. PhD thesis, University of British Columbia, Vancouver, B.C. 104 pages.
Wittmer, H.U., R.N.M. Ahrens and B.N. McLellan. 2010. Viability of mountain caribou in British Columbia, Canada: Effects of habitat change and population density. Biological Conservation 143: 86–93. Available here.
Wittmer, H.U., B.N. McLellan, D.R. Seip, J.A. Young, T.A. Kinley, G.S. Watts and D. Hamilton. 2005a. Population dynamics of the endangered mountain ecotype of woodland caribou (Rangifer tarandus caribou) in British Columbia, Canada. Canadian Journal of Zoology 83: 407–418.
Wittmer, H.U., A.R.E. Sinclair and B.N. McLellan. 2005b. The role of predation in the decline and extirpation of woodland caribou. Oecologia 144: 257–267.
Wittmer, H.U., B.N. McLellan and F.W. Hovey. 2006. Factors influencing variation in site fidelity of woodland caribou (Rangifer tarandus caribou) in southeastern British Columbia. Canadian Journal of Zoology 84: 537–545.
Wittmer, H.U., B.N. McLellan, R. Serrouya and C.D. Apps. 2007. Changes in landscape composition influence the decline of a threatened woodland caribou population. Journal of Animal Ecology 76: 568–579.